Dr. Nathan Horrell

Lecturer of Biology

Nathan HorrellEducation

  • B.S., Behavioral Neuroscience, Centre College
  • Ph.D., Neuroscience, University of California, Riverside


Electrophysiology, behavioral endocrinology, parental care

Research Interest

My research at the University of California, Riverside focused on the neuroendocrinology of parental care, specifically paternal care in mammals. Using a biparental mammalian model species, the California mouse, I investigated the neural plasticity that underlies their shift in infant-directed behavior, from indifferent or aggressive to robustly paternal, that occurs in virgin male California mice when they become fathers. My approach to teaching uses active learnings and emphasizes the utility of course material.


  • Horrell, N.D., Saltzman, W., Hickmott, P. (2019). Plasticity of paternity: effects of fatherhood on synaptic, intrinsic and morphological characteristics of neurons in the medial preoptic area of male California mice. Behavioural Brain Research, 365, 89-102.
  • Perea-Rodriguez, J. P., de Jong, T. R., Kung, E., Horrell, N. D., Saltzman W. (2019). Consequences of placentophagia by adult virgin male California mice (Peromyscus californicus). Behavioural Processes, 166, 103889.
  • Horrell N.D., Hickmott, P., Saltzman, W. (2018). Neural regulation of paternal behavior in mammals: sensory, neuroendocrine, and experiential influences on the paternal brain. Grattan, D. & Coolen, L., Eds., Current Topics in Behavioral Neuroscience. New York, NY: Springer.
  • Horrell, N.D., Perea-Rodriguez, J. P., Harris, B. N., Saltzman, W. (2017). Effects of repeated pup exposure on behavioral, neural, and adrenocortical responses to pups in male California mice (Peromyscus californicus). Hormones and Behavior, 90, 56-63
  • Saltzman, W., Harris, B.N., De Jong, T.R., Perea-Rodriguez, J.P., Horrell, N.D., Zhao, M., Andrew, J.R. (2017). Paternal care in biparental rodents: Intra- and inter-individual variation. Integrative and Comparative Biology, 57, 589–602.

Contact Dr. Nathan Horrell

(478) 301-2836